  {"id":10524,"date":"2022-03-21T15:25:31","date_gmt":"2022-03-21T19:25:31","guid":{"rendered":"http:\/\/149.4.100.129\/academics\/bio\/?page_id=10524"},"modified":"2025-10-29T18:10:47","modified_gmt":"2025-10-29T22:10:47","slug":"doctorate","status":"publish","type":"page","link":"https:\/\/www.qc.cuny.edu\/academics\/bio\/doctorate\/","title":{"rendered":"Doctorate"},"content":{"rendered":"<p>[et_pb_section fb_built=&#8221;1&#8243; _builder_version=&#8221;4.16&#8243; background_color=&#8221;rgba(0,0,0,0)&#8221; custom_padding=&#8221;16px|||||&#8221; global_colors_info=&#8221;{}&#8221;][et_pb_row _builder_version=&#8221;4.16&#8243; _module_preset=&#8221;default&#8221; global_colors_info=&#8221;{}&#8221;][et_pb_column type=&#8221;4_4&#8243; _builder_version=&#8221;4.16&#8243; _module_preset=&#8221;default&#8221; global_colors_info=&#8221;{}&#8221;][dsm_text_divider header=&#8221;Doctoral Studies&#8221; text_alignment=&#8221;left&#8221; color=&#8221;#E71939&#8243; divider_position=&#8221;flex-end&#8221; divider_weight=&#8221;5px&#8221; admin_label=&#8221;Doctoral Studies&#8221; _builder_version=&#8221;4.16&#8243; _module_preset=&#8221;default&#8221; header_font=&#8221;Open Sans|600|||||||&#8221; header_text_color=&#8221;#000000&#8243; header_font_size=&#8221;30px&#8221; global_colors_info=&#8221;{}&#8221;][\/dsm_text_divider][\/et_pb_column][\/et_pb_row][et_pb_row column_structure=&#8221;3_5,2_5&#8243; _builder_version=&#8221;4.16&#8243; _module_preset=&#8221;default&#8221; global_colors_info=&#8221;{}&#8221;][et_pb_column type=&#8221;3_5&#8243; _builder_version=&#8221;4.16&#8243; _module_preset=&#8221;default&#8221; global_colors_info=&#8221;{}&#8221;][et_pb_text admin_label=&#8221;Description&#8221; _builder_version=&#8221;4.27.4&#8243; text_font=&#8221;Open Sans||||||||&#8221; text_font_size=&#8221;16px&#8221; header_4_font=&#8221;Open Sans|600|||on||||&#8221; text_orientation_tablet=&#8221;left&#8221; text_orientation_phone=&#8221;&#8221; text_orientation_last_edited=&#8221;on|desktop&#8221; global_colors_info=&#8221;{}&#8221;]<\/p>\n<p>Although the various CUNY senior colleges do not grant doctoral degrees, PhD students who enroll at the <a href=\"http:\/\/www.gc.cuny.edu\/\" target=\"_blank\" rel=\"noopener\">CUNY Graduate Center<\/a> can conduct their dissertation research under the advisement of QC Biology faculty members.<\/p>\n<p>Most PhD students in QC Biology labs are enrolled in one of four <a href=\"https:\/\/www.gc.cuny.edu\/biology\" target=\"_blank\" rel=\"noopener\">PhD Biology<\/a> subprograms: Evolution, Ecology, and Behavior (EEB); Molecular, Cellular, and Developmental Biology (MCD); Neuroscience; and Plant Sciences. However, some enrol in other PhD Programs, notably Psychology (Subprograms in Behavioral and Cognitive Neuroscience; and Cognition and Comparative Psychology), Biochemistry, or Earth and Environmental Sciences.<\/p>\n<p>Please refer to the <a href=\"https:\/\/www.gc.cuny.edu\/academic-bulletin\" target=\"_blank\" rel=\"noopener\">Bulletin of the Graduate Center<\/a> for information on applying, financial aid, and courses.<\/p>\n<p>The QC Biology Department has excellent research\u00a0facilities, including the <a href=\"https:\/\/www.qc.cuny.edu\/academics\/bio\/core-facilities\/\" target=\"_blank\" rel=\"noopener self\">Core Facility for Imaging, Cellular and Molecular Biology<\/a>, well-equipped research labs, a greenhouse, animal facilities, cold-room and environmental control units, marine and fresh-water aquaria, scanning and transmission electron microscopes, and computer access. Our doctoral faculty do <a href=\"https:\/\/www.qc.cuny.edu\/academics\/bio\/research\/\" target=\"_blank\" rel=\"noopener\">research across a wide range of topics<\/a> and in a wide variety of systems around the world, as exemplified by the work of Dr Sebastian Alvarado (see photo and caption at right).<\/p>\n<p>&nbsp;<\/p>\n<p>[\/et_pb_text][\/et_pb_column][et_pb_column type=&#8221;2_5&#8243; _builder_version=&#8221;4.16&#8243; _module_preset=&#8221;default&#8221; global_colors_info=&#8221;{}&#8221;][et_pb_image src=&#8221;https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-content\/uploads\/sites\/117\/2025\/10\/Doctoral_studies_img_446wide.png&#8221; alt=&#8221;PhD student Matthew Pero\u0161 and Thomas (a local field assistant) are pictured in a boat on Lake Tanganyika in Zambia, They are photographing cichlids.&#8221; title_text=&#8221;Doctoral_studies_img_446wide&#8221; _builder_version=&#8221;4.27.4&#8243; _module_preset=&#8221;default&#8221; global_colors_info=&#8221;{}&#8221;][\/et_pb_image][et_pb_text _builder_version=&#8221;4.27.4&#8243; _module_preset=&#8221;default&#8221; custom_margin=&#8221;-17px|||||&#8221; global_colors_info=&#8221;{}&#8221;]<\/p>\n<p><span data-olk-copy-source=\"MessageBody\">Matthew Pero\u0161, at left, is doing his PhD with Dr Alvarado, studying Africa&#8217;s endemic cichlids. Here, he and his local field assistant, Thomas (at right) are photographing <\/span><span style=\"background-color: #ffffff;font-size: 14px\"><em>Astatotilapia burtoni<\/em> in Z<\/span><span style=\"font-size: 14px;background-color: #ffffff\">ambia&#8217;s L<\/span><span style=\"font-size: 14px;background-color: #ffffff\">ake Tanganyika <\/span><span style=\"font-size: 14px;background-color: #ffffff\">to investigate how cichlids&#8217; colors change with seasons.<\/span><\/p>\n<p>[\/et_pb_text][\/et_pb_column][\/et_pb_row][et_pb_row _builder_version=&#8221;4.16&#8243; _module_preset=&#8221;default&#8221; custom_padding=&#8221;0px|||||&#8221; global_colors_info=&#8221;{}&#8221;][et_pb_column type=&#8221;4_4&#8243; _builder_version=&#8221;4.16&#8243; _module_preset=&#8221;default&#8221; global_colors_info=&#8221;{}&#8221;][et_pb_text admin_label=&#8221;Description&#8221; _builder_version=&#8221;4.27.4&#8243; text_font=&#8221;Open Sans||||||||&#8221; text_font_size=&#8221;16px&#8221; header_4_font=&#8221;Open Sans|600|||on||||&#8221; min_height=&#8221;1247.5px&#8221; custom_margin=&#8221;-35px|||||&#8221; text_orientation_tablet=&#8221;left&#8221; text_orientation_phone=&#8221;&#8221; text_orientation_last_edited=&#8221;on|desktop&#8221; global_colors_info=&#8221;{}&#8221;]<\/p>\n<p>Opportunities are enhanced by affiliation with other New York City (NYC) institutions and cooperative efforts with other CUNY divisions and the American Museum of Natural History (AMNH). PhD-level courses can be taken at the Graduate Center, QC or any other CUNY college affiliated with the Graduate Center (<a href=\"http:\/\/www.brooklyn.cuny.edu\/\" target=\"_blank\" rel=\"noopener\">Brooklyn College<\/a>, <a href=\"http:\/\/www.ccny.cuny.edu\/\" target=\"_blank\" rel=\"noopener\">City College<\/a>, <a href=\"http:\/\/www.csi.cuny.edu\/\" target=\"_blank\" rel=\"noopener\">College of Staten Island<\/a>, <a href=\"http:\/\/www.hunter.cuny.edu\/\" target=\"_blank\" rel=\"noopener\">Hunter College<\/a>, <a href=\"http:\/\/www.lehman.cuny.edu\/\" target=\"_blank\" rel=\"noopener\">Lehman College<\/a>). Four other institutions in the NYC area are associated with the CUNY PhD program in Biology, including the <a href=\"http:\/\/www.amnh.org\/\" target=\"_blank\" rel=\"noopener\">AMNH<\/a>, <a href=\"http:\/\/www.bbg.org\/\" target=\"_blank\" rel=\"noopener\">Brooklyn Botanic Garden<\/a>, the <a href=\"https:\/\/opwdd.ny.gov\/institute-basic-research-developmental-disabilities\" target=\"_blank\" rel=\"noopener\">Institute for Basic Research in Developmental Disabilities<\/a>, and <a href=\"http:\/\/www.nybg.org\/\" target=\"_blank\" rel=\"noopener\">The New York Botanical Garden<\/a>. Our faculty and doctoral students collaborate and interact with researchers at academic and other research institutions in NYC, elsewhere in the United States and abroad. ºì¶¹ÊÓÆµ is near many habitats in urban, forest, freshwater, brackish, coastal and marine ecosystems.<\/p>\n<p>All CUNY PhD students receive guaranteed financial support (stipend and full tuition) for five years. This financial support is covered by the <a href=\"http:\/\/www.gc.cuny.edu\/\" target=\"_blank\" rel=\"noopener\">CUNY Graduate Center<\/a> in year 1 (when PhD students mainly take classes) and by the faculty mentor and campus in subsequent years. Consult other relevant PhD programs for their policies.<\/p>\n<p>Below are links to CUNY PhD subprograms that QC Biology faculty are affiliated with.<\/p>\n<p><a href=\"https:\/\/www.gc.cuny.edu\/biology\/subprograms\/ecology-evolutionary-biology-and-behavior\" target=\"_blank\" rel=\"noopener self\">Ecology\/Evolution\/Behavior<\/a><br \/>Sebastian Alvarado<br \/>Mitchell Baker<br \/>Joanna Coleman<br \/>John Dennehy<br \/>David Lahti<br \/>John Waldman<\/p>\n<p><a href=\"https:\/\/www.gc.cuny.edu\/biology\/subprograms\/molecular-cellular-and-developmental-biology\" target=\"_blank\" rel=\"noopener\">Molecular\/Cell\/Developmental Biology<\/a><br \/>Sebastian Alvarado<br \/>John Dennehy<br \/>Tim Dubuc<br \/>Alicia Mel\u00e9ndez<br \/>Cathy Savage-Dunn<br \/>Maral Tajerian<br \/>Daniel Weinstein<\/p>\n<p><a href=\"https:\/\/www.gc.cuny.edu\/biology\/subprograms\/neuroscience\" target=\"_blank\" rel=\"noopener\">Neuroscience Subprogram<\/a><br \/>Sebastian Alvarado<br \/>Maral Tajerian<\/p>\n<p><a href=\"https:\/\/www.gc.cuny.edu\/biochemistry\" target=\"_blank\" rel=\"noopener\">Biochemistry PhD Program<\/a><br \/>Karl Fath<br \/>Alicia Mel\u00e9ndez<br \/>Daniel Weinstein<\/p>\n<p><a href=\"https:\/\/www.gc.cuny.edu\/psychology\" target=\"_blank\" rel=\"noopener\">Psychology PhD Program<\/a><br \/>David Lahti<\/p>\n<p>[\/et_pb_text][\/et_pb_column][\/et_pb_row][\/et_pb_section]<\/p>\n","protected":false},"excerpt":{"rendered":"<p><div class=\"et_pb_module dsm_text_divider dsm_text_divider_0\">\n\t\t\t\t\n\t\t\t\t\n\t\t\t\t\n\t\t\t\t\n\t\t\t\t\n\t\t\t\t\n\t\t\t\t<div class=\"et_pb_module_inner\">\n\t\t\t\t\t<div class=\"dsm-text-divider-wrapper dsm-text-divider-align-left et_pb_bg_layout_light\">\n\t\t\t\t\n\t\t\t\t\n\t\t\t\t<div class=\"dsm-text-divider-before dsm-divider\"><\/div>\n\t\t\t\t<h3 class=\"dsm-text-divider-header et_pb_module_header\"><span>Doctoral Studies<\/span><\/h3>\n\t\t\t\t<div class=\"dsm-text-divider-after dsm-divider\"><\/div>\n\t\t\t<\/div>\n\t\t\t\t<\/div>\n\t\t\t<\/div>Although the various CUNY senior colleges do not grant doctoral degrees, PhD students who enroll at the CUNY Graduate Center can conduct their dissertation research under the advisement of QC Biology faculty members. Most PhD students in QC Biology labs [&hellip;]<\/p>\n","protected":false},"author":2,"featured_media":0,"parent":0,"menu_order":0,"comment_status":"closed","ping_status":"closed","template":"","meta":{"_et_pb_use_builder":"on","_et_pb_old_content":"","_et_gb_content_width":"","inline_featured_image":false,"footnotes":""},"page_category":[],"wf_page_folders":[134],"class_list":["post-10524","page","type-page","status-publish","hentry"],"_links":{"self":[{"href":"https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-json\/wp\/v2\/pages\/10524","targetHints":{"allow":["GET"]}}],"collection":[{"href":"https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-json\/wp\/v2\/pages"}],"about":[{"href":"https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-json\/wp\/v2\/types\/page"}],"author":[{"embeddable":true,"href":"https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-json\/wp\/v2\/users\/2"}],"replies":[{"embeddable":true,"href":"https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-json\/wp\/v2\/comments?post=10524"}],"version-history":[{"count":0,"href":"https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-json\/wp\/v2\/pages\/10524\/revisions"}],"wp:attachment":[{"href":"https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-json\/wp\/v2\/media?parent=10524"}],"wp:term":[{"taxonomy":"page_category","embeddable":true,"href":"https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-json\/wp\/v2\/page_category?post=10524"},{"taxonomy":"wf_page_folders","embeddable":true,"href":"https:\/\/www.qc.cuny.edu\/academics\/bio\/wp-json\/wp\/v2\/wf_page_folders?post=10524"}],"curies":[{"name":"wp","href":"https:\/\/api.w.org\/{rel}","templated":true}]}}